Habitats
Native soft scales
Some native species are "host specific", meaning they can live only on particular plants, for example, the button scales [Lecanochiton species] are only found on pōhutukawa and rātā. Other native soft scales can be found on several different plants, but usually in just one habitat.
Beech (Nothofagus)
Two different soft scale insects are found on Nothofagus species in beech forest:
- Conical beech scale Crystallotesta fagi (Maskell)
- Glassy beech scale Crystallotesta neofagi Henderson & Hodgson
![Conical beech scale [Crystallotesta fagi] (Maskell) Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Crystallotesta_fagi__FillMaxWzE1LDE1XQ.jpg)
![[Crystallotesta fagi]. L: a young female fills her clear waxy test and her greenish colour shows through. R: an old female her body shrunk to the anterior end of her white glassy wax test [lower end in picture] after producing all her crawlers. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Crystallotesta_fagi_stages__FillMaxWzE1LDE1XQ.jpg)
![Glassy beech scale [Crystallotesta neofagi] Henderson & Hodgson Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Crystallotesta_neofagi__FillMaxWzE1LDE1XQ.jpg)
Kānuka and Mānuka
Kānuka [Kunzea ericoides] and Mānuka [Leptospermum scoparium]
There are four different soft scale insects found in this habitat:
- Glassy mānuka scale Crystallotesta leptospermi (Maskell)
- Small ornate scale Crystallotesta ornatella Henderson & Hodgson
- Thumbnail scale Plumichiton pollicinus Henderson & Hodgson
- Knobbly scale Umbonichiton bullatus Henderson & Hodgson
![[Crystallotesta leptospermi]. L: young adult female, her colour pattern mimics the leaf bud scales of the host plant kanuka. R: old female, her body has shrunk to a small dark lump at the head [head facing down the twig] after producing her young. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Crystallotesta_leptospermi_2__FillMaxWzE1LDE1XQ.jpg)
![[Crystallotesta ornatella] L: young female; R: old female Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Crystallotesta_ornatella-v2__FillMaxWzE1LDE1XQ.jpg)
![[Plumichiton pollicinus]. L: young adult female, the upper middle part is of curled white wax layers, the flatter sides show her body colour through transparent wax plates. R: mature female. Females always have their head-ends facing down the twig. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Plumichiton_pollicinus__FillMaxWzE1LDE1XQ.jpg)
![[Umbonichiton bullatus]. Young adult female. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Umbonichiton_bullatus__FillMaxWzE1LDE1XQ.jpg)
Lowland
Mixed broadleaf forest types, other than podocarps and kānuka.
There are 17 different soft scale insects found in mixed broadleaf forest:
- Translucent scale Aphenochiton pubens Henderson & Hodgson
- Filmy scale Aphenochiton subtilis Henderson & Hodgson
- Ornate scale Crystallotesta ornata (Maskell)
- Turtleshell scale Ctenochiton chelyon Henderson & Hodgson
- Spotted sixpenny scale Ctenochiton paraviridis Henderson & Hodgson
- Toronia scale Ctenochiton toru Henderson & Hodgson
- Sixpenny scale Ctenochiton viridis Maskell
- Peppercorn scale Epelidochiton piperis (Maskell)
- Limpet scale Inglisia patella Maskell
- Glassy coprosma scale Kalasiris depressa (Maskell)
- Fringed scale Kalasiris perforata (Maskell)
- Golden plumed scale Plumichiton flavus (Maskell)
- Black plumed scale Plumichiton elaeocarpi (Maskell)
- Rough scale Umbonichiton hymenantherae (Maskell)
![[Aphenochiton pubens] - adult female. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Aphenochiton_pubens__FillMaxWzE1LDE1XQ.jpg)
![[Aphenochiton subtilis]. Adult female. The rows of waxy plates covering the insect body show up particularly well because this photo was taken with cross-lighting. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Aphenochiton_subtilis__FillMaxWzE1LDE1XQ.jpg)
![[Crystallotesta ornata] Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Crystallotesta_ornata__FillMaxWzE1LDE1XQ.jpg)
![[Ctenochiton chelyon]. Two adult females together and several of their crawlers (young scales) walking around them. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Ctenochiton_chelyon__FillMaxWzE1LDE1XQ.jpg)
![[Ctenochiton paraviridis]. L: a young adult female. R: a mature female with several new crawlers nearby on the leaf. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Ctenochiton_paraviridis__FillMaxWzE1LDE1XQ.jpg)
![[Ctenochiton toru]. A mature female. She has caused the leaf to curl on its edge. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Ctenochiton_toru__FillMaxWzE1LDE1XQ.jpg)
![[Ctenochiton viridis]. A mature female with a new crawler walking away on the leaf. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Ctenochiton_viridis__FillMaxWzE1LDE1XQ.jpg)
![[Epelidochiton piperis]. L: Four young adult females (various yellow - green - greyish-brown colours). R: a mature female (dark greyand the smaller white scale is an empty male test (cover) the adult male has already emerged. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Epelidochiton_piperis__FillMaxWzE1LDE1XQ.jpg)
![[Inglisia patella]. Young adult female. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Inglisia_patella__FillMaxWzE1LDE1XQ.jpg)
![[Kalasiris depressa]. L: young adult female. R: mature adult female. Her body (greenish colour) has shrunk into the front half of her waxy test. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Kalasiris_depressa__FillMaxWzE1LDE1XQ.jpg)
![[Kalasiris perforata]. L: young adult female. R: adult female secreting new wax on the pattern lines of her test. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Kalasiris_perforata__FillMaxWzE1LDE1XQ.jpg)
![[Plumichiton flavus]. Adult female. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Plumichiton_flavus__FillMaxWzE1LDE1XQ.jpg)
![[Plumichiton elaeocarpi]. L: young adult female. R: mature adult female. She has a hardened black skin and only the remains of a wax test: a little on the top and a worn down marginal fringe. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Plumichiton_elaeocarpi___FillMaxWzE1LDE1XQ.jpg)
![[Umbonichiton hymenantherae]. Young adult female. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Umbonichiton_hymenantherae__FillMaxWzE1LDE1XQ.jpg)
![[Aphenochiton kamahi] - adult female. The yellow part in the middle is her unborn crawlers, the small orange-brown spot her anal plates (where she gets rid of honeydew); the outer part is nearly transparent. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Aphenochiton_kamahi__FillMaxWzE1LDE1XQ.jpg)
![[Plumichiton nikau]. An adult female. She is convex with a thick glassy wax test. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Plumichiton_nikau__FillMaxWzE1LDE1XQ.jpg)
![[Pounamococcus cuneatus]. L: young adult female. R: mature adult female. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Pounamococcus_cuneatus__FillMaxWzE1LDE1XQ.jpg)
Near seashore
Sometimes inland But always found on Pōhutukawa and rātā [Metrosideros species]
three button scales are found here:
- Pōhutukawa button scale Lecanochiton actites Henderson & Hodgson
- Rātā button scale Lecanochiton metrosideri Maskell
- Small button scale Lecanochiton scutellaris Henderson & Hodgson
![[Crystallotesta ornata]. L: a young adult female. R: a mature female; both are facing down the twig. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Crystallotesta_ornata_stages__FillMaxWzE1LDE1XQ.jpg)
![[Poropeza dacrydii]. L: 3rd-instar female (yellow) and young adult female (reddish colour), on the underside of bark taken from a totara tree. R: mature female with a crawler (orangey colour) walking over her. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Poropeza_dacrydii__FillMaxWzE1LDE1XQ.jpg)
![[Umbonichiton adelus]. Young adult female. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Umbonichiton_adelus__FillMaxWzE1LDE1XQ.jpg)
Exotic soft scales
Some of the exotic species have very wide host plant ranges (they are polyphagous). In areas disturbed by human activity, e.g., bush margins, they can even be found on some native plants, and two of them, soft brown scale and hemispherical scale, have spread a little bit further into native forest.
Gardens
Eight different types of soft scale insects are found on shrubs, trees and ferns in areas modified by humans (gardens and home orchards):
Chinese wax scale Ceroplastes sinensis Del Guercio
Soft brown scale Coccus hesperidum Linnaeus
Long brown scale Coccus longulus (Douglas)
Nigra scale Parasaissetia nigra (Nietner)
Cottony hydrangea scale Pulvinaria hydrangeae Steinweden
Ice plant scale Pulvinaria mesembryanthemi (Vallot)
Hemispherical scale Saissetia coffeae (Walker)
Olive scale or black scale Saissetia oleae (Olivier)
![[Ceroplastes sinensis] L: immatures: 2nd-instars (white) and 3rd-instars (pink). R: adult females; they have "buttons" and stripes of a different kind of wax that is left over from the immature stages. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Ceroplastes_sinensi__FillMaxWzE1LDE1XQ.jpg)
![[Coccus hesperidum]. L: immature and young adult females. R: two adult females and immature females. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Coccus_hesperidum__FillMaxWzE1LDE1XQ.jpg)
![[Coccus longulus] - young adult female, on stem of citrus. She is facing head to the right and her two eye spots are small black dots, one on either side. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Coccus_longulus__FillMaxWzE1LDE1XQ.jpg)
![[Parasaissetia nigra]. Two adult females and some settled crawlers (pale yellow-green). Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Parasaissetia_nigra__FillMaxWzE1LDE1XQ.jpg)
![[Saissetia coffeae]. L: immature and young adult females. R: mature female. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Saissetia_coffeae__FillMaxWzE1LDE1XQ.jpg)
![[Saissetia oleae]. L: immature female. R: mature female. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Saissetia_oleae__FillMaxWzE1LDE1XQ.jpg)
![[Pulvinaria hydrangeae]. Adult female, with her long white eggsac and the young crawlers hatching out of it. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Pulvinaria_hydrangeae__FillMaxWzE1LDE1XQ.jpg)
![[Pulvinaria mesembryanthemi] Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Pulvinaria_mesembryanthemi__FillMaxWzE1LDE1XQ.jpg)
Orchards
Five different soft scale insects are found on citrus, stonefruits and feijoas in orchards:
Soft wax scale Ceroplastes destructor Newstead
Chinese wax scale Ceroplastes sinensis Del Guercio
Brown scale Parthenolacanium corni (Bouche)
Cottony vine scale Pulvinaria vitis (Linnaeus)
Black scale or olive scale Saissetia oleae (Olivier)
![[Ceroplastes destructor] - adult females. The one on the left is undamaged and shows the stripes of harder wax on its side. The female on the right's soft wax has been slightly squished. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Ceroplastes_destructor__FillMaxWzE1LDE1XQ.jpg)
![[Ceroplastes sinensis] L: immatures: 2nd-instars (white) and 3rd-instars (pink). R: adult females; they have "buttons" and stripes of a different kind of wax that is left over from the immature stages. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Ceroplastes_sinensi__FillMaxWzE1LDE1XQ.jpg)
![[Parthenolecanium corni]. Adult females on a small branch of cherry. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Parthenolecanium_corni__FillMaxWzE1LDE1XQ.jpg)
![[Pulvinaria vitis]. L: young adult female. R: mature female with eggsac (white and fluffy). As the eggs are laid into the soft wax, the female's body is lifted up until it is about at right angles to the branch. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Pulvinaria_vitis__FillMaxWzE1LDE1XQ.jpg)
![[Saissetia oleae]. L: immature female. R: mature female. Image](/assets/Discover-Our-Research/Biodiversity/Plants-fungi-arthropods-bacteria/Invertebrate-Systematics/scale-insects/Saissetia_oleae__FillMaxWzE1LDE1XQ.jpg)